The 27-year-old singer opened up the show with a performance of her song “Applause” and later met up with the group One Direction after getting off stage.
“Macklemore is so amazing!!! What a night!” Gaga tweeted after the singer spoke out in support of gay rights while accepting an award.
FYI: Zayn is wearing a Saint Laurent leather jacket.
Lady Gaga Gives Pep Talk to One Direction